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Emmer

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(Redirected from Triticum dicoccoides)
For other uses, see Emmer (disambiguation).

Emmer wheat
Spikes (ears) of cultivated emmer wheat
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Monocots
Clade: Commelinids
Order: Poales
Family: Poaceae
Subfamily: Pooideae
Genus: Triticum
Species:
T. dicoccum
Binomial name
Triticum dicoccum
Synonyms[4]
  • Spelta amylea (Ser.) Ser.
  • Triticum amyleum Ser.
  • Triticum armeniacum (Stolet.) Nevski
  • Triticum arras Hochst.
  • Triticum atratum Host
  • Triticum cienfuegos Lag.
  • Triticum dicoccum Schrank.[3]
  • Triticum dicoccoides
  • Triticum farrum Bayle-Bar.
  • Triticum gaertnerianum Lag.
  • Triticum immaturatum Flaksb. nom. inval.
  • Triticum ispahanicum Heslot
  • Triticum karamyschevii Nevski
  • Triticum maturatum Flaksb. nom. inval.
  • Triticum palaecocolchicum (Menabde) L.B. Cai
  • Triticum palaeocolchicum Menabde
  • Triticum subspontaneum (Tzvelev) Czerep.
  • Triticum tricoccum Schübl.
  • Triticum volgense (Flaksb.) Nevski
Look up emmer in Wiktionary, the free dictionary.

Emmer wheat or hulled wheat[3] is a type of awned wheat. Emmer is a tetraploid (4n = 4x = 28 chromosomes).[5] The domesticated types are Triticum turgidum subsp. dicoccum and T. t. conv. durum. The wild plant is called T. t. subsp. dicoccoides. The principal difference between the wild and the domestic forms is that the ripened seed head of the wild plant shatters and scatters the seed onto the ground, while in the domesticated emmer, the seed head remains intact, thus making it easier for people to harvest the grain.[6]

Along with einkorn wheat, emmer was one of the first crops domesticated in the Near East. It was widely cultivated in the ancient world, but is now a relict crop in mountainous regions of Europe and Asia.

Emmer is considered a type of farro food, especially in Italy.[3]

Etymology

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Emmer is first attested in 1908 in English as a loanword from German Emmer, variant of Amelkorn, from amel, 'starch', likely from Latin amylum, itself borrowing from Ancient Greek amylon.[7]

Taxonomy

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Strong similarities in morphology and genetics show that wild emmer (T. dicoccoides Koern.) is the wild ancestor and a crop wild relative of domesticated emmer. Because wild and domesticated emmer are interfertile with other tetraploid wheats, some taxonomists consider all tetraploid wheats to belong to one species, T. turgidum. Under this scheme, the two forms are recognized at subspecies level, thus T. t. subsp. dicoccoides and T. t. subsp. dicoccum. Either naming system is equally valid; the latter lays more emphasis on genetic similarities.[citation needed]

For a wider discussion, see Wheat § Genetics and breeding and Wheat taxonomy

Wild emmer

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Wild emmer grows wild in the Near East. It is a tetraploid wheat formed by the hybridization of two diploid wild grasses, Triticum urartu, closely related to wild einkorn (T. boeoticum), and an as yet unidentified Aegilops species related to A. searsii or A. speltoides.[citation needed]

Botanists Körnicke and Aaronsohn in the late 19th-century were the first to describe the wild emmer (Triticum dicoccoides) native to Palestine and adjacent countries.[8][9][10] Earlier, in 1864, the Austrian botanist Kotschy had collected specimens of the same wild emmer, without signifying where he had collected them.[11]

Although cultivated in ancient Egypt, wild emmer has not been cultivated for human consumption in recent history,[9] perhaps owing to the difficulty with which the chaff is separated from the seed kernels, formerly requiring the spikes to be pounded with mortar and pestle.[12]

The wild emmer is distinguished from T. vulgare, with its tougher ear rhachis and the beards releasing the grains easily, by their ear rhachis that are brittle when ripe and their firmly fitting beards.[9] The wild emmer grows to a height of 50–70 cm (20–28 in), and bears an elongated spike measuring 10–15 cm (3.9–5.9 in), with long, protruding awns extending upwards.[11]

Morphology

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Grains of a cultivated type
Cultivated type

Like einkorn and spelt wheats, emmer is a hulled wheat, meaning it has strong glumes (husks) that enclose the grains, and a semibrittle rachis. On threshing, a hulled wheat spike breaks up into spikelets that require milling or pounding to release the grains from the glumes.[citation needed]

Wild emmer wheat spikelets effectively self-cultivate by propelling themselves mechanically into soils with their awns. During a period of increased humidity during the night, the awns of the spikelet become erect and draw together, and in the process push the grain into the soil. During the daytime, the humidity drops and the awns slacken back again; however, fine silica hairs on the awns act as hooks in the soil and prevent the spikelets from backing out. During the course of alternating stages of daytime drying and nighttime humidity, the awns' pumping movements, which resemble a swimming frog kick, will drill the spikelet 25 millimetres (1 inch) or more into the soil.[13]

History

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Wild emmer is native to the Fertile Crescent of the Middle East, growing in the grass and woodland of hill country from modern-day Israel to Iran. The origin of wild emmer has been suggested, without universal agreement among scholars, to be the Karaca Dağ mountain region of southeastern Turkey. In 1906, Aaron Aaronsohn's discovery of wild emmer wheat growing in Rosh Pinna (Israel) created a stir in the botanical world.[14] Emmer wheat has been found in archaeological excavations and ancient tombs. Emmer was collected from the wild and eaten by hunter gatherers for thousands of years before its domestication. Grains of wild emmer discovered at Ohalo II had a radiocarbon dating of 17,000 BC and at the Pre-Pottery Neolithic A (PPNA) site of Netiv Hagdud are 10,000–9,400 years old.[15]

The location of the earliest site of emmer domestication is still unclear and under debate.[16] Some of the earliest sites with possible indirect evidence for emmer domestication during the Early Pre-Pottery Neolithic B include Tell Aswad, Çayönü, Cafer Höyük, Aşıklı Höyük, Kissonerga-Mylouthkia [de] and Shillourokambos.[16] Definitive evidence for the full domestication of emmer wheat is not found until the Middle Pre-Pottery Neolithic B (10,200 to 9,500 BP), at sites such as Beidha, Tell Ghoraifé, Tell es-Sultan (Jericho), Abu Hureyra, Tell Halula, Tell Aswad and Cafer Höyük.[16]

Wild wheat (Triticum turgidum subsp. dicoccoides) growing in Israel

Emmer is found in a large number of Neolithic sites scattered around the fertile crescent. From its earliest days of cultivation, emmer was a more prominent crop than its cereal contemporaries and competitors, einkorn wheat and barley.[17] Small quantities of emmer are present during Period 1 at Mehrgharh on the Indian subcontinent, showing that emmer was already cultivated there by 7000–5000 BC.[18]

In the Near East, in southern Mesopotamia in particular, cultivation of emmer wheat began to decline in the Early Bronze Age, from about 3000 BC, and barley became the standard cereal crop. This has been related to increased salinization of irrigated alluvial soils, of which barley is more tolerant,[19] although this study has been challenged.[20] Emmer had a special place in ancient Egypt, where it was the main wheat cultivated in Pharaonic times, although cultivated einkorn wheat was grown in great abundance during the Third Dynasty, and large quantities of it were found preserved, along with cultivated emmer wheat and barleys, in the subterranean chambers beneath the Step Pyramid at Saqqara.[21] Neighbouring countries also cultivated einkorn, durum and common wheat.[22] In the absence of any obvious functional explanation, the greater prevalence of emmer wheat in the diet of ancient Egypt may simply reflect a marked culinary or cultural preference, or may reflect growing conditions having changed after the Third Dynasty. Emmer and barley were the primary ingredients in ancient Egyptian bread and beer. Emmer recovered from the Phoenician settlement at Volubilis[23] (in present-day Morocco) has been dated to the middle of the first millennium BC.

Emmer wheat may be one of the five species of grain which have a special status in Judaism. One of these species, referred to as kusmin (Aramaic) or kūsmīn (Hebrew: כוסמין), may be either emmer or spelt (emmer and spelt are easily confused with each other). However, it is fairly certain that spelt did not grow in ancient Israel, and emmer was probably a significant crop until the end of the Iron Age. References to emmer in Greek and Latin texts are traditionally translated as "spelt", even though spelt was not common in the Classical world until very late in its history.[citation needed]

In northeastern Europe, emmer (in addition to einkorn and barley) was one of the most important cereal species and this importance can be seen to increase from 3400 BC onward. Pliny the Elder notes that although emmer was called far in his time formerly it was called adoreum (or 'glory'), providing an etymology explaining that emmer had been held in glory (N.H. 18.3), and later in the same book he describes its role in sacrifices.[citation needed]

Cultivation

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Emmer in NRW, Germany, affected by lodging

Today emmer is primarily a relict crop in mountainous areas. Its value lies in its ability to give good yields on poor soils, and its resistance to fungal diseases such as stem rust that are prevalent in wet areas. Emmer is grown in Armenia, Morocco, Spain (Asturias), the Carpathian mountains on the border of Czechia and Slovakia, Albania, Turkey, Switzerland, Germany, Greece and Italy. It is also grown in the U.S. as a specialty product. A traditional food plant in Ethiopia, this relatively little-known grain has potential to improve nutrition, boost food security, foster rural development and support sustainable landcare.[24]

In Italy, uniquely, emmer cultivation is well established and even expanding. In the mountainous Garfagnana area of Tuscany emmer (known as farro) is grown by farmers as an IGP (Indicazione Geografica Protetta) product, with its geographic identity protected by law. Production is certified by a co-operative body, the Consorzio Produttori Farro della Garfagnana. IGP-certified farro is widely available in health food shops across Europe, and even in some British supermarkets. The demand for Italian farro has led to competition from non-certified farro, grown in lowland areas and often consisting of a different wheat species, spelt (T. spelta).[citation needed]

Food uses

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Minestra di farro [it] is a typical dish in Tuscany, Italy

Emmer's main use is as a human food, though it is also used for animal feed. Ethnographic evidence from Turkey and other emmer-growing areas suggests that emmer makes good bread (judged by the taste and texture standards of traditional bread), and this is supported by evidence of its widespread consumption as bread in ancient Egypt.[25] Emmer bread is available in Switzerland[26] and the Netherlands. The largest Dutch supermarket chain in the Netherlands, Albert Heijn, sells emmer bread as part of their luxury bread line.

In Armenian cuisine, emmer pilaf is cooked in sunflower oil. The pre-cooked emmer is fried in oil and then sautéed onions are added. In the second version, boiled water and emmer are added to the sautéed onion and then cooked until tender.[27] In Armenia, emmer porridge with lamb called "kashovi" is widespread in the Shirak region, while lean kashovi is known in all regions of Armenia. Chopped lamb is added to cold water then boiled for 10 minutes, emmer and sautéed onion are added and cooked over a low heat until tender.[28]

In Italy, whole emmer grains can be easily found in most supermarkets and groceries, emmer bread (pane di farro) can be found in bakeries in some areas, while in Tuscany emmer has traditionally been consumed as whole grains in soup. Emmer has also been used in beer production.[29]

In parts of India, emmer wheat (in Maharashtra called खपली गहू, transcription "khapalī gahū", meaning "crusty wheat") is grown as a drought- and stress-resistant wheat variety with some work ongoing to improve yields, as a result of increased interest in this variety due to possible value for diabetics.[30]

As with all varieties and hybrids of wheat,[31] emmer is unsuitable for people with gluten-related disorders, such as celiac disease, non-celiac gluten sensitivity and people with wheat allergies, among others.[32][33]

Genome

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Avni et al., 2017 provides a genome.[34]

Notes

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  1. ^ Zair, W. & Magos Brehm, J. (2017). "Triticum turgidum Wild Emmer". IUCN Red List of Threatened Species. 2017: e.T71786841A71786853. doi:10.2305/IUCN.UK.2017-3.RLTS.T71786841A71786853.en.
  2. ^ "Tropicos.org". Archived from the original on 3 March 2016. Retrieved 30 June 2014.
  3. ^ a b c "Triticum turgidum subsp. dicoccon". Germplasm Resources Information Network. Agricultural Research Service, United States Department of Agriculture. Retrieved 11 December 2017.
  4. ^ "The Plant List: A Working List of All Plant Species". Retrieved June 30, 2014.
  5. ^ "Complex Bread Wheat Genome Cracked". Nat Geo Food. 17 July 2017. Archived from the original on July 17, 2014.
  6. ^ Weiss, Ehud and Zohary, Daniel (October 2011), "The Neolithic Southwest Asian Founder Crops", Current Anthropology, Vo 52, Supplement 4, p. S240
  7. ^ "emmer". Online Etymology Dictionary. Retrieved 2011-08-10.
  8. ^ Zohary, M. (1998). David Heller (ed.). A New Analytical Flora of Israel (in Hebrew) (2 ed.). Tel Aviv: Am Oved. p. 562. OCLC 916628298. (first edition 1976)
  9. ^ a b c Dalman, Gustaf (2020). Nadia Abdulhadi-Sukhtian (ed.). Work and Customs in Israel, volume II. Vol. 2 (Agriculture). Translated by Robert Schick. Ramallah: Dar Al Nasher. pp. 288–289. ISBN 978-9950-385-84-9.
  10. ^ Aaronsohn, A. (1910). Agricultural and botanical explorations in Israel. Washington, D.C.: United States Department of Agriculture. pp. 42-ff., 46-ff. OCLC 631059153.
  11. ^ a b Shmida, Avi (2005). MAPA's Dictionary of Plants and Flowers in Israel (in Hebrew). Tel-Aviv: MAPA. p. 393. OCLC 716569354.
  12. ^ Amar, Zohar (2015). Flora and Fauna in Maimonides' Teachings (in Hebrew). Kfar Darom. pp. 96–97. OCLC 783455868.{{cite book}}: CS1 maint: location missing publisher (link)
  13. ^ Elbaum, Rivka; Zaltzman, Liron; Burgert, Ingo; Fratzl, Peter (2007). "The Role of Wheat Awns in the Seed Dispersal Unit". Science. 316 (5826): 884–886. Bibcode:2007Sci...316..884E. doi:10.1126/science.1140097. PMID 17495170. S2CID 42898148.
  14. ^ Molecular Genetic Maps in Wild Emmer Wheat, Triticum dicoccoides: Genome-Wide Coverage, Massive Negative Interference, and Putative Quasi-Linkage
  15. ^ Zohary & Hopf 2000, p. 46
  16. ^ a b c Weide, Alexander (2015). "On the Identification of Domesticated Emmer Wheat, Triticum turgidum subsp. dicoccum (Poaceae), in the Aceramic Neolithic of the Fertile Crescent". Archäologische Informationen [de].
  17. ^ Weiss and Zohary, pp. S240-S242
  18. ^ Possehl, Gregory. "The Indus Civilization: An Introduction to Environmental, Subsistence, and Cultural History: (2003)
  19. ^ Jacobsen & Adams 1958
  20. ^ Powell, M. A. (1985) Salt, seed, and yields in Sumerian agriculture. A critique of the theory of progressive salinization. Zeitschrift für Assyriologie und Vorderasiatische Archäologie 75, 7–38.
  21. ^ Jean-Phillipe Lauer, Laurent Taeckholm and E. Aberg, 'Les Plantes Decouvertes dans les Souterrains de l'Enceinte du Roi Zoser a Saqqarah' in Bulletin de l'Institut d'Egypte, Vol. XXXII, 1949–50, pp. 121–157, and see Plate IV for photo of ears of both wheats recovered from beneath the pyramid.
  22. ^ Zohary & Hopf 2000, pp. 50f
  23. ^ C. Michael Hogan. 2008. Volubilis: Ancient settlement in Morocco, The Megalithic Portal, ed. Andy Burnham
  24. ^ National Research Council (1996-02-14). "Other Cultivated Grains". Lost Crops of Africa: Volume I: Grains. Lost Crops of Africa. Vol. 1. National Academies Press. ISBN 978-0-309-04990-0. Retrieved 2008-07-18.
  25. ^ Hulled wheats. Proceedings of the First International Workshop on Hulled Wheats. Promoting the conservation and use of underutilized and neglected crops 4. Edited by S. Padulosi, K. Hammer, and J. Heller, 1996. Rome: International Plant Genetic Resources Institute.[permanent dead link‍]
  26. ^ "Renaissance alter Brotgetreidesorten – swissinfo" (in German). Swissinfo.ch. Retrieved 2010-11-13.[permanent dead link‍]
  27. ^ G. Khanbekyan. A book about the national Armenian food. For housewives. Yerevan, Armgiz, 1950. p. 80
  28. ^ G. Khanbekyan. A book about the national Armenian food. For housewives, Yerevan, Armgiz, 1950, p. 82
  29. ^ Samuel, Delwen. 1996. Archeology of Ancient Egyptian Beer. Journal of the American Society of Brewing Chemists 54(1): 3–12
  30. ^ Urs, Anil (23 February 2011). "Medicinal traits found in 'Khapli' wheat". @businessline. Retrieved 2019-08-16.
  31. ^ Fasano A, Sapone A, Zevallos V, Schuppan D (May 2015). "Nonceliac gluten sensitivity". Gastroenterology. 148 (6): 1195–204. doi:10.1053/j.gastro.2014.12.049. PMID 25583468.
  32. ^ Tovoli F, Masi C, Guidetti E, Negrini G, Paterini P, Bolondi L (Mar 16, 2015). "Clinical and diagnostic aspects of gluten related disorders". World J Clin Cases. 3 (3): 275–84. doi:10.12998/wjcc.v3.i3.275. PMC 4360499. PMID 25789300.
  33. ^ "Grains in Relation to Celiac (Coeliac) Disease". Wheat.pw.usda.gov. Archived from the original on 2010-10-18. Retrieved 2010-11-13.
  34. ^
    Steensels, Jan; Gallone, Brigida; Voordeckers, Karin; Verstrepen, Kevin (2019). "Domestication of Industrial Microbes". Current Biology. 29 (10). Cell Press: R381–R393. Bibcode:2019CBio...29.R381S. doi:10.1016/j.cub.2019.04.025. ISSN 0960-9822. PMID 31112692. S2CID 159040577.
    Fernie, Alisdair R.; Yan, Jianbing (2019). "De Novo Domestication: An Alternative Route toward New Crops for the Future". Molecular Plant. 12 (5). Cell Press: 615–631. doi:10.1016/j.molp.2019.03.016. ISSN 1674-2052. PMID 30999078. S2CID 121615993.
    These reviews cite this research.
    Avni, Raz; Nave, Moran; Barad, Omer; Baruch, Kobi; Twardziok, Sven; Gundlach, Heidrun; Hale, Iago; Mascher, Martin; Spannagl, Manuel; Wiebe, Krystalee; Jordan, Katherine; Golan, Guy; Deek, Jasline; Ben-Zvi, Batsheva; Ben-Zvi, Gil; Himmelbach, Axel; MacLachlan, Ron; Sharpe, Andrew; Fritz, Allan; Ben-David, Roi; Budak, Hikmet; Fahima, Tzion; Korol, Abraham; Faris, Justin; Hernandez, Alvaro; Mikel, Mark; Levy, Avraham; Steffenson, Brian; Maccaferri, Marco; Tuberosa, Roberto; Cattivelli, Luigi; Faccioli, Primetta; Ceriotti, Aldo; Kashkush, Khalil; Pourkheirandish, Mohammad; Komatsuda, Takao; Eilam, Tamar; Sela, Hanan; Sharon, Amir; Ohad, Nir; Chamovitz, Daniel; Mayer, Klaus; Stein, Nils; Ronen, Gil; Peleg, Zvi; Pozniak, Curtis; Akhunov, Eduard; Distelfeld, Assaf (2017). "Wild emmer genome architecture and diversity elucidate wheat evolution and domestication". Science. 357 (6346). American Association for the Advancement of Science: 93–97. Bibcode:2017Sci...357...93A. doi:10.1126/science.aan0032. ISSN 0036-8075. PMID 28684525. S2CID 19943904.

References

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